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I acknowledge the value of many thoughtful discussions with Melanie Cooper over the last few years on the conceptual foundations of chemistry and biology and how they might be taught more effectively. I also thank the organizers of, and participants in, the ASBMB/National Science Foundation (NSF)–sponsored Conceptual Assessment in Biochemistry and Molecular Biology meeting (March 2010), particularly Ellis Bell, Duane Sears, and Robin Wright, for their inspiration, as well as the University of Colorado Boulder/CU Teach Discipline-based Education Research (DBER) group for providing an ongoing forum for the discussion of a wide range of science education issues. My work in biology education has been supported by NSF grants DUE 0405007 and DUE 0816692 and can be accessed through the beSocratic.colorado.edu and Bioliteracy.colorado.edu websites.