We thank Jeff Hardin and the Department of Integrative Biology at the University of Wisconsin–Madison for funding through the Michael Guyer Post-Doctoral Fellowship. We also thank students in the Biocore program and the UW Zoological Museum for lending S. vulgaris specimens. We are grateful for interview transcription by Maddie Batzli and many conversations with colleagues, including Michelle Harris, Emily Jobe, and Robin Forbes-Lorman, whose insights and questions have been invaluable.